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Artie Abrams (Kevin McHale) has taken charge of directing the school musical, West Side Story, and he tells the two leads—Rachel (Lea Michele), playing Maria, and Blaine (Darren Criss), playing Tony—that they are not conveying enough emotion in their rendition of "Tonight". He questions whether they can convincingly portray the roles if they are still virgins. Later, his co-director Coach Beiste (Dot-Marie Jones) confesses to him that she is attracted to football recruiter Cooter Menkins (Eric Bruskotter), who is at McKinley to scout potential players for Ohio State, though she is sure he would never consider her. Artie thinks otherwise, and acting on his advice, Cooter arranges a date with her.

Blaine and Kurt (Chris Colfer) discuss their decision to postpone sex. Blaine goes to Dalton Academy to invite the Warblers to see him in West Side Story and arrives as they are performing "Uptown Girl". Afterward, new Warbler Sebastian Smythe (Grant Gustin) makes a play for Blaine, intercut with Santana (Naya Rivera) and Rachel singing "A Boy Like That" from the musical. They later meet at a coffee shop, and Kurt arrives as Blaine is telling Sebastian that he already has a boyfriend. Sebastian convinces them to go with him to the local gay bar and supplies fake IDs. While Blaine and Sebastian are dancing, Kurt is surprised to see Dave Karofsky (Max Adler), who has transferred to another school. When Blaine and Kurt leave, Blaine is drunk and aroused, and urges Kurt to have sex with him in the back seat of the car; Kurt refuses, and Blaine walks home.

Rachel lets Finn (Cory Monteith) know that she is interested in having sex with him, but he balks when she admits her reason is to be a better Maria. Later, she asks advice from the other girls in the show. While Santana and Quinn (Dianna Agron) both urge her not to have sex with Finn, Tina (Jenna Ushkowitz) reveals that she and Mike (Harry Shum, Jr.) had sex over the summer and says how wonderful it was with the boy she loved—her words are intercut with Rachel and Santana singing "I Have a Love" in rehearsal. Mike's father (Keong Sim) confronts him about his participation in the musical, and Mike tells him he wants to be a professional dancer, not a doctor. His father disowns him.

Cooter recruits Shane (LaMarcus Tinker) for Ohio State, but not Finn, who is despondent about his future. Rachel comforts him, and promises that she will help him find a new future. Back at school, Blaine apologizes to Kurt for getting drunk, and says he cares nothing for Sebastian. Kurt proposes that they go over to Blaine's house for the night.

On opening night, Artie is assailed by self-doubt, but he is thanked by the cast for his leadership, and he thanks them for trusting him. They perform "America", which gets a standing ovation. Blaine and Rachel, waiting to go on and still virgins, are afraid they will not convey the necessary emotion, but Rachel reminds Blaine that they both have found their soulmates in Finn and Kurt, just like Maria and Tony had with each other. As they sing "One Hand, One Heart" on stage, they are also shown in scenes involving their first sexual encounters with their true soulmates.

Information from Wikipedia